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Effects of Cross and Self Fertilisation in the Vegetable Kingdom by Charles Darwin
page 298 of 636 (46%)
other exotic plants, which have been self-fertilised for many
generations. In such cases a cross between two individuals does no good;
nor does it in any case, unless the individuals differ in general
constitution, either from so-called spontaneous variation, or from their
progenitors having been subjected to different conditions. I believe
that this is the true explanation in the present instance, because, as
we shall immediately see, the offspring of plants, which did not profit
at all by being crossed with a plant of the same stock, profited to an
extraordinary degree by a cross with a slightly different sub-variety.

THE EFFECTS OF A CROSS WITH A FRESH STOCK.

I procured some seed of N. tabacum from Kew and raised some plants,
which formed a slightly different sub-variety from my former plants; as
the flowers were a shade pinker, the leaves a little more pointed, and
the plants not quite so tall. Therefore the advantage in height which
the seedlings gained by this cross cannot be attributed to direct
inheritance. Two of the plants of the third self-fertilised generation,
growing in Pots 2 and 5 in Table 6/87, which exceeded in height their
crossed opponents (as did their parents in a still higher degree) were
fertilised with pollen from the Kew plants, that is, by a fresh stock.
The seedlings thus raised may be called the Kew-crossed. Some other
flowers on the same two plants were fertilised with their own pollen,
and the seedlings thus raised from the fourth self-fertilised
generation. The crossed capsules produced by the plant in Pot 2, Table
6/87, were plainly less fine than the self-fertilised capsules on the
same plant. In Pot 5 the one finest capsule was also a self-fertilised
one; but the seeds produced by the two crossed capsules together
exceeded in number those produced by the two self-fertilised capsules on
the same plant. Therefore as far as the flowers on the parent-plants are
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