Insectivorous Plants by Charles Darwin
page 18 of 532 (03%)
page 18 of 532 (03%)
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cells there is an inner one of differently shaped ones, likewise filled
with purple fluid, but of a slightly different tint, and differently affected by chloride of gold. These two layers are sometimes well seen when a gland has been crushed or boiled in caustic potash. According to Dr. Warming, there is still another layer of much more elongated cells, as shown in the accompanying section (fig. 3) copied from his work; but these cells were not seen by Nitschke, nor by me. In the centre there is a group of elongated, cylindrical cells of unequal lengths, bluntly pointed at their upper ends, truncated or rounded at their lower ends, closely pressed together, and remarkable from being surrounded by a spiral line, which can be separated as a distinct fibre. These latter cells are filled with limpid fluid, which after long immersion in alcohol deposits much brown matter. I presume that they are actually connected with the spiral vessels which run up the tentacles, for on several occasions the latter were seen to divide into two or three excessively thin branches, which could be traced close up to the spiriferous cells. Their development has been described by Dr. Warming. Cells of the same kind have been observed in other plants, as I hear from Dr. Hooker, and were seen by me in the margins of the leaves of Pinguicula. Whatever their function may be, they are not necessary for the secretion of a digestive fluid, or for absorption, or for the communication of a motor impulse to other parts of the leaf, as we may infer from the structure of the glands in some other genera of the Droseraceae. The extreme marginal tentacles differ slightly from the others. Their bases are broader, and besides their own vessels, they receive a fine branch from those which enter the tentacles on each side. Their glands are much elongated, and lie embedded on the upper surface of the |
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