Facts and Arguments for Darwin by Fritz Muller
page 102 of 127 (80%)
page 102 of 127 (80%)
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larvae; and thus, in the Isopoda, the historically newest pair of feet
is produced later than all the rest. In the Copepoda this formation of new segments and limbs, gradually advancing from before backwards, is more perfectly preserved than in any of the higher Crustacea.* (* It is well known that, in many cases, even in adult animals the last segment of the middle-body, or some of its last segments, either want their limbs or are themselves deficient (Entoniscus Porcellanae male, Leucifer, etc.). This might be due to the animals having separated from the common stem before these limbs were formed at all. But in those cases with which I am best acquainted, it seems to me more probable that the limbs have been subsequently lost again. That these particular limbs and segments are more easily lost than others is explained by the circumstance that, as the youngest, they have been less firmly fixed by long-continued inheritance. ("Mr. Dana believes, that in ordinary Crustaceans, the abortion of the segments with their appendages almost always takes place at the posterior end of the cephalothorax."--Darwin, Balanidae, page 111.)) The original development of the Malacostraca starting from the Nauplius, or the lowest free-living grade with which we are acquainted in the class of Crustacea, is now-a-days nearly effaced in the majority of them. That this extinction has actually taken place in the way already deduced as a direct consequence from Darwin's theory, will be the more easily demonstrated, the more this process is still included in the course of life, and the less completely it is already worn out. We may hope to obtain the most striking examples in the still unknown developmental history of the various Schizopoda, Peneidae, and, indeed, of the Macrura in general. At present the multifarious Zoea-forms appear to be particularly instructive. Almost all the peculiarities by which they depart from the primitive form of the Zoea of Peneus (Figures 29, |
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