Facts and Arguments for Darwin by Fritz Muller
page 60 of 127 (47%)
page 60 of 127 (47%)
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Caprella.) a peculiar structure makes its appearance very early on the
anterior part of the back, by which the embryo is attached to the "inner egg-membrane," and which has been called the "micropylar apparatus," but improperly as it seems to me.* (* Little as a name may actually affect the facts, we ought certainly to confine the name "micropyle" to canals of the egg-membrane, which serve for the entrance of the semen. But the outer egg-membrane passes over the "micropylar apparatus" of the Amphipoda without any perforation, according to Meissner's and La Valette's own statements; it appears never to be present before fecundation, attains its greatest development at a subsequent period of the ovular life, and the delicate canals which penetrate it do not even seem to be always present, indeed it seems to belong to the embryo rather than to the egg-membrane. I have never been able to convince myself that the so-called "inner egg-membrane" is really of this nature, and not perhaps the earliest larva skin, not formed until after impregnation, as might be supposed with reference to Ligia, Cassidina and Philoscia.) It will remind us of the union of the young Isopoda with the larval membrane and of the unpaired "adherent organ" on the nape of the Cladocera, which is remarkably developed in Evadne and persists throughout life; but in Daphnia pulex, according to Leydig, although present in the young animals, disappears without leaving a trace in the adults. The young animal, whilst still in the egg, acquires the full number of its segments and limbs. In cases where segments are amalgamated together, such as the last two segments of the thorax in Dulichia, the last abdominal segments and the tail in Gammarus ambulans and Corophium dentatum, n. sp., and the last abdominal segments and the tail in Brachyscelus,* or where one or more segments are deficient, as in Dulichia and the Caprellae, we find the same fusion and the same |
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